Residency at University of Florida
- 1982 – 1984, Small Animal Internal Medicine
Past Professional Activities at University of Pennsylvania
- 1984 – 1985, Instructor in Small Animal Medicine
- 1985 – 1990, Assistant Professor of Medicine and Medical Genetics
- 1989 – 2018, Director of the Metabolic Genetics Laboratory - PennGen
- 1989 – present, Secondary Appointment as Privatdozent/Professor at University of Zürich
- 1989 – 2009, Medical Director of Transfusion Medicine Center
- 1990 – 1995, Associate Professor of Medicine and Medical Genetics
- 1990 – present, Secondary Appointment as Professor of Hematology, School of Medicine
- 1995 – 2007, Professor of Medical Genetics and Chief of Medical Genetics
- 1997 – present, Charlotte Newton Sheppard Endowed Professor of Medicine
- 1998 – present, Director of PennGen Laboratories
Honors and Awards
- 1984 – Phi Zeta National Veterinary Honor Society
- 2002 – Scientific Lifetime Achievement Award from the World Small Animal Veterinary Association
- 2002 – Transfusion Medicine Academic Award
- 2004 – John E. McCoy Award from Washington State University for excellence in scientific studies
- 2007 – International Bourgelat Award from British Small Animal Veterinary Association for outstanding contributions to the field of medicine
- 2008 – 2018 WSAVA Hereditary Disease Committee Chair
- 2015 – AVMA Excellence in Feline Research Award for outstanding contributions to feline medicine
- 2018 – Ramsey Distinguished Lecture Award from Iowa State University for clinical research
Current Research/Clinical Interests
His clinical and research expertise and interests are in clinical hematologic disorders, transfusion medicine, hereditary diseases and genetic predispositions in companion animals. They are reflected in >250 original research publications as well as many more reviews and scientific abstracts. He is a frequently invited speaker at national and international conferences.
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